Speed and memory
In our performance index the GeForce RTX 5090 scores 130 and the GeForce RTX 4080 Super scores 73, which makes the GeForce RTX 5090 about 78% faster. That is a gap you will notice: roughly one step up in quality setting, or the difference between 60 and 80 fps.
The GeForce RTX 5090 has 32 GB of video memory and the GeForce RTX 4080 Super has 16 GB. That difference matters at 1440p and in new games at High textures, and it can outweigh a small speed advantage as the card ages.
In our estimates for popular games at 1080p on High, the biggest gap is in Call of Duty (274 against 163 fps), and the smallest is in ELDEN RING (60 against 60 fps). Games that are light enough to run above 200 fps on both cards show little difference at all.